The monophyly of the Imicola complex, a natural species complex within subgenus C. subgen. Avaritia Fox of the biting midge genus Culicoides Latreille, is supported using morphological and molecular analyses. A diagnosis for the group along with comparative redescriptions of the male and female of the species represented in Australasia, C. brevitarsis Kieffer and C. nudipalpis Delfinado and a description of C. asiatica Bellis sp. nov., are presented together with keys for their specific determination and molecular support for their status.
Bellis, G., Dyce, A., Gopurenko, D., Yanase, T., Garros, C., Labuschagne, K., & Mitchell, A. (2014). Revision of the Culicoides (Avaritia) Imicola complex Khamala & Kettle (Diptera : Ceratopogonidae) from the Australasian region. Zootaxa, 3768(4), 401-427. https://doi.org/10.11646/zootaxa.3768.4.1